The National Rifle Association’s Telegenic Warrior

Ann Coulter found her voice as a Cassandra for the right in the early 2000s. In the Trump era, Ms. Loesch has found hers as a telegenic warrior for the N.R.A. She shares some characteristics with President Trump, said Megyn Kelly, the NBC talk show host who used to have her on as a guest on Fox News when Ms. Kelly had a show there. “She is compelling television,” she said.

Even Ms. Kelly, a friend, has noticed that Ms. Loesch has embraced a more barbed tone the past year. “On my show, she was never a bomb thrower,” Ms. Kelly said. “We tried to veer away from that.” But public conversation has grown more cacophonous since Mr. Trump’s election. And unlike Ms. Coulter in her heyday, Ms. Loesch faces an ever-expanding field of conservative pundits willing to do anything to entertain followers on Twitter, in their jostle for the seats at Fox’s table.

“I’ve been told I’m a whore for the N.R.A.,” Ms. Loesch said. “I’m a prostitute for the N.R.A. But I believe so strongly in the natural right to bear arms. I feel so passionately about that.”

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Ms. Loesch on her show in 2014, broadcasting from CPAC, an annual conference for American conservatives.

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Michael Mathes/Agence France-Presse — Getty Images

Dismissal only firms her resolve. “I’m feeling like I am doing what I want to do,” she said, “and what I feel fulfilled in doing.”

Wonder Woman

For as long as Ms. Loesch can remember, the world has been a scary place. She was born Dana Eaton in 1978 in Hematite, Mo., a small community about 40 miles south of St. Louis. Her parents divorced when she was in kindergarten and she has not spoken to her father in more than a decade. Her mother worked three jobs after they moved to nearby Festus, Mo., bringing home bags of granola bars after her shift at a local granola factory. Ms. Loesch said she spent school-day afternoons at home alone or with an aunt.

“Now, you could say it was like a meth ’hood,” Ms. Loesch said of her old neighborhood. “People were fighting. It was kind of crazy. It wasn’t the most stable of childhoods.”

Ms. Loesch mostly found refuge at her grandparents’ home in Annapolis, Mo., a town of about 450 people nestled in the rural Ozarks. Her family voted for Democrats. Her grandfather hunted deer and raccoon. Despite episodes of violence, Ms. Loesch idealized summers in Annapolis in “Flyover Country.” In the book, she recalled her grandfather standing on the porch one night with a shotgun in his hands. Ms. Loesch’s aunt had just arrived; her estranged husband had threatened to kill her.

“Looking back,” Ms. Loesch said, “I think I always wanted to know that I was safe.”

In the late 1990s, Ms. Loesch attended Webster University in suburban St. Louis, studying journalism, but dropped out after she got pregnant. In 2000, she married Chris Loesch, the baby’s father: a musician and son of a preacher who now manages her career. They attend weekly services at the Church of Christ. And on Halloween they give a party that harks back to more pleasant childhood memories. “Halloween was big with her mom,” said Leigh Wambsganns, a friend. Last October Ms. Loesch said she dressed as Wonder Woman.

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Ms. Loesch and her husband, Chris, listen to basic firearm safety instructions.

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Allison V. Smith for The New York Times

As a new parent in St. Louis, she blogged about motherhood and began her long-running radio program, “The Dana Show.” She became disillusioned with Democratic politics, though, in the wake of President Clinton’s affair with Monica Lewinsky. She helped found the St. Louis Tea Party, organizing protests and speaking at rallies. And, in 2010, she joined Andrew Breitbart at his website, one of several new voices railing against establishment politics and media bias.

After Mr. Breitbart died in 2012, Ms. Loesch clashed with Steve Bannon, the former Trump ally who had been named executive chair of Breitbart News. She sued to get out of her contract. In 2014, she moved to Dallas to work for The Blaze.

Ms. Loesch’s particular brand of attack proved unpalatable to mainstream audiences. In 2012, CNN, which had hired her as a political commentator, distanced itself from comments she made on her radio show supporting a group of Marines who urinated on dead Taliban soldiers. She claims the reporting of her comments was “disingenuous.” A year later, she was banned from the now defunct “Piers Morgan Live” after getting into a Twitter fight with the host. “I thought we made up,” she said. “But then we started fighting again.”

And, in 2016, Ms. Loesch expressed personal ire at Kayleigh McEnany, now the spokeswoman for the Republican National Committee. Ms. McEnany had chided a friend of Ms. Loesch’s for not supporting Mr. Trump, then a Republican candidate, during a CNN segment. “Babycakes, this was more than just going on television and flashing your pearly whites and your flat chest, red dress, over-sprayed bleach blond hair,” Ms. Loesch told radio listeners.

This time, it seemed, she had gone too far. Ms. McEnany was undergoing treatment for a genetic predisposition to breast cancer. Both Ms. Loesch and Ms. McEnany declined to discuss the matter.

Home on the Range

Ms. Loesch’s performance for the N.R.A. has had more polish. Last April, she was featured in a recruiting ad for the organization called “Violence of Lies.” Scenes of street violence and protests flashed onscreen as Ms. Loesch called for citizens to fight media bias and liberal politicians with the “clenched fist of truth.” (The N.R.A. also released an ad in 2017 aimed at The New York Times, claiming media bias.)

According to Andrew Arulanandam, the N.R.A.’s managing director of public affairs, the message was “inspired” by the chief executive, Wayne LaPierre Jr., whose association donated $30 million in campaign support to Mr. Trump. According to news reports, the N.R.A. may have a connection to the investigation of Russia’s involvement in the election. “Violence of Lies” has been viewed nearly three million times on YouTube alone.

“Dana comes across clearly to our members and gun owners,” Mr. Arulanandam said.

But even some in those groups found the video unnecessarily, even dangerously, incendiary. An online petition was circulated, demanding Facebook remove it. On Twitter, DeRay Mckesson, a Black Lives Matter activist, called the video “an open call to violence to protect white supremacy.” In response, Ms. Loesch challenged him to come on her show and “tell me to my face I’m a racist.” He didn’t, saying Ms. Loesch seeks to use opponents as a foil.

“I don’t believe that she was really talking to me,” he said in an interview. “She was using me to rile up the base.” Last week Ms. Loesch said she had no interest in speaking with Mr. Mckesson.

Ms. Loesch’s boss, Mr. LaPierre, has a history of inflammatory rhetoric at the N.R.A., which has five million members. In 1995, he was forced to apologize after President George H. W. Bush canceled his N.R.A. membership in protest. The N.R.A. had sent out a fund-raising letter calling law enforcement “jackbooted government thugs” who threatened to hurt Americans. “That is what they do,” said Representative Kathleen Rice, a Democrat of New York, who has sparred with Ms. Loesch, also on Twitter. “The N.R.A says their members are under attack.”

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